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A girl named Penny and a dog named Bolt star on a hit television series called Bolt in which the titular character has various superpowers and must constantly thwart the evil plans of the nefarious Doctor Calico. To gain a more realistic performance, the TV show's producers have deceived Bolt his entire life, arranging the filming in such a way that Bolt believes the television show is real and he really has superpowers. After filming completes for the latest episode, Bolt escapes from his on-set trailer mistakenly believing Penny has been kidnapped by the television villain. He attempts to break through a window, knocking himself unconscious as he falls into a box of styrofoam peanuts. With no one aware Bolt is in the box, it is shipped from Hollywood to New York City. When Bolt arrives in New York, he meets a sour-attitude cat named Mittens, who was declawed and abandoned by her owners. The pair travel to Ohio, where they then meet a TV-obsessed hamster named Rhino who just doesn't seem to leave his hamster ball. The three embark on a quest to find Penny again.
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